Table of Contents
Study Area Map A Note on Accompanying Video Robert K. Landis Foreword Jane Goodall Preface Douglas W. Smith, Daniel R. Stahler, and Daniel R. MacNultyPart 1 History and Reintroduction 1 Historical and Ecological Context for Wolf Recovery Douglas W. Smith, Daniel R. Stahler, Daniel R. MacNulty, and Lee H. Whittlesey Box 1.1 Wolf History and Surveys in Yellowstone National Park
John Weaver 2 How Wolves Returned to Yellowstone
Steven H. Fritts, Rebecca J. Watters, Edward E. Bangs, Douglas W. Smith, and Michael K. Phillips
Box 2.1 To Reintroduce or Not to Reintroduce, That Is the Question Diane Boyd Guest Essay: Why Are Yellowstone Wolves Important?
L. David MechPart 2 Behavioral and Population Ecology 3 Essential Biology of the Wolf: Foundations and Advances Daniel R. MacNulty, Daniel R. Stahler, Tim Coulson, and Douglas W. Smith 4 Ecology of Family Dynamics in Yellowstone Wolf Packs
Daniel R. Stahler, Douglas W. Smith, Kira A. Cassidy, Erin E. Stahler, Matthew C. Metz, Rick McIntyre, and Daniel R. MacNulty Box 4.1 Naming Wolf Packs
Daniel R. Stahler 5 Territoriality and Competition between Wolf Packs
Kira A. Cassidy, Douglas W. Smith, Daniel R. Stahler, Daniel R. MacNulty, Erin E. Stahler, and Matthew C. Metz
Box 5.1 Auditory Profile: The Howl of the Wolf John B. Theberge and Mary T. Theberge 6 Population Dynamics and Demography
Douglas W. Smith, Kira A. Cassidy, Daniel R. Stahler, Daniel R. MacNulty, Quinn Harrison, Ben Balmford, Erin E. Stahler, Ellen E. Brandell, and Tim Coulson Guest Essay: Yellowstone Wolves Are Important Because They Changed Science
Rolf O. Peterson and Trevor S. PetersonPart 3 Genetics and Disease 7 Yellowstone Wolves at the Frontiers of Genetic Research
Daniel R. Stahler, Bridgett M. vonHoldt, Elizabeth Heppenheimer, and Robert K. Wayne 8 The K Locus: Rise of the Black Wolf
Rena M. Schweizer, Daniel R. Stahler, Daniel R. MacNulty, Tim Coulson, Phil Hedrick, Rachel Johnston, Kira A. Cassidy, Bridgett M. vonHoldt, and Robert K. Wayne 9 Infectious Diseases in Yellowstone’s Wolves
Ellen E. Brandell, Emily S. Almberg, Paul C. Cross, Andrew P. Dobson, Douglas W. Smith, and Peter J. Hudson Guest Essay: Why Are Yellowstone Wolves Important? A European Perspective
Olof LibergPart 4 Wolf-Prey Relationships 10 How We Study Wolf-Prey Relationships
Douglas W. Smith, Matthew C. Metz, Daniel R. Stahler, and Daniel R. MacNulty
Box 10.1 Nine-Three-Alpha Douglas W. Smith
Box 10.2 The Bone Collectors Ky Koitzsch and Lisa Koitzsch 11 Limits to Wolf Predatory Performance
Daniel R. MacNulty, Daniel R. Stahler, and Douglas W. Smith
Box 11.1 Tougher Times for Yellowstone Wolves Reflected in Tooth Wear and Fracture Blaire Van Valkenburgh 12 What Wolves Eat and Why
Matthew C. Metz, Mark Hebblewhite, Douglas W. Smith, Daniel R. Stahler, Daniel R. MacNulty, Aimee Tallian, and John A. Vucetich
Box 12.1 Bison in Wood Buffalo National Park L. N. Carbyn 13 Wolf Predation on Elk in a Multi-Prey Environment
Matthew C. Metz, Douglas W. Smith, Daniel R. Stahler, Daniel R. MacNulty, and Mark Hebblewhite
Box 13.1 Generalizing Wolf-Prey Dynamics across Systems: Yellowstone, Banff, and Isle Royale Mark Hebblewhite
Box 13.2 The Predator’s Perspective: Biomass of Prey Matthew C. Metz Box 13.3 Lessons from Denali National Park: Stability in Predator-Prey Dynamics Is a Pause on the Way to Somewhere Else Layne Adams 14 Population Dynamics of Northern Yellowstone Elk after Wolf Reintroduction
Daniel R. MacNulty, Daniel R. Stahler, Travis Wyman, Joel Ruprecht, Lacy M. Smith, Michel T. Kohl, and Douglas W. Smith
Box 14.1 Wolves and Elk in the Madison Headwaters Robert A. Garrott, P. J. White, Claire Gower, Matthew S. Becker, Shana Drimal, Ken L. Hamlin, and Fred G. R. Watson
Box 14.2 Ecology of Fear Daniel R. Stahler and Daniel R. MacNulty Guest Essay: The Value of Yellowstone’s Wolves? The Power of Choice
Michael K. PhillipsPart 5 Ecosystem Effects and Species Interactions 15 Indirect Effects of Carnivore Restoration on Vegetation
Rolf O. Peterson, Robert L. Beschta, David J. Cooper, N. Thompson Hobbs, Danielle Bilyeu Johnston, Eric J. Larsen, Kristin N. Marshall, Luke E. Painter, William J. Ripple, Joshua R. Rose, Douglas W. Smith, and Evan C. Wolf
Box 15.1 Long-Term Trends in Beaver, Moose, and Willow Status in the Southern Portion of the Absaroka-Beartooth Wilderness Daniel B. Tyers 16 Competition and Coexistence among Yellowstone’s Meat Eaters
Daniel R. Stahler, Christopher C. Wilmers, Aimee Tallian, Colby B. Anton, Matthew C. Metz, Toni K. Ruth, Douglas W. Smith, Kerry A. Gunther, and Daniel R. MacNulty Guest Essay: Old Dogs Taught Old Lessons
Paul C. PaquetPart 6 Conservation, Management, and the Human Experience 17 Wolves and Humans in Yellowstone
Douglas W. Smith, Daniel R. Stahler, Rick McIntyre, Erin E. Stahler, and Kira A. Cassidy 18 The Wolf Watchers
Nathan Varley, Rick McIntyre, and James Halfpenny
Box 18.1 Bob Landis’s Yellowstone Wolves Documentaries 000 Box 18.2 Seeing Wolves Robert Hayes 19 Conservation and Management: A Way Forward
Douglas W. Smith, P. J. White, Daniel R. Stahler, Rebecca J. Watters, Kira A. Cassidy, Adrian Wydeven, Jim Hammill, and David E. Hallac Guest Essay: Making Better Sense of Wolves
Susan G. Clark
Afterword Rebecca J. Watters, Douglas W. Smith, Daniel R. Stahler, and Daniel R. MacNulty Acknowledgments Appendix: Species Names Used in the Text Literature Cited List of Contributors Author Index Subject Index