Swifty the Arctic Fox (Jeremy Renner) spends his days working away in Arctic Blast Delivery Service's mailroom. However, he secretly longs to become a Top Dog, one of the company's prized husky couriers. When an unusual package passes through the service, Swifty decides to steal a sled and deliver the package to its mystery destination to prove his skill. Along the way, he unwittingly stumbles upon a troublesome plot concocted by the sinister Otto Von Walrus (John Cleese). It is up to Swifty and his rag-tag team of pals to put a stop to Otto's scheme once and for all.